* Jørgensen, P. M. & C. Ulloa Ulloa. 1994. Seed plants of the high Andes of Ecuador—A checklist. AAU Rep. 34: 1-443
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* Standley, P. C. & J. A. Steyermark. 1946. Rosaceae. En: Standley, P.C. & J.A. Steyermark (eds.), Flora of Guatemala - Part IV. Fieldiana, Bot. 24(4): 432-484